If you look at the volume for the days in question when the short positions could not be covered, there are millions of shares traded. 8 million, followed by 5 million, then 7 million. Those three day's totals together are greater than the float. I suspect that uncovered short sales had been probably numbered in several million shares; so, to take losses, someone has to eat several hundred thousand dollars to possibly a million or so. That is a lot of cheese to eat, and covering in a bullish situation only feeds the run, thus making losses greater. Damned if they do, and damned if they don't. IMHO
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